Eva Hedlund

Om mig
Publikationer
CV
Populärvetenskap och samverkan

Forskningsbeskrivning

Mer information finns tillgänglig på den engelska profilsidan [1]
samt på lab hemsidan [2]
[1] http://ki.se/en/people/evahed
[2] http://ki.se/en/neuro/hedlund-laboratory

Artiklar

Journal article: EUROPEAN NEUROPSYCHOPHARMACOLOGY. 2024;78:54-63

Substance P, NPY, CCK and their receptors in five brain regions in major depressive disorder with transcriptomic analysis of locus coeruleus neurons.

Barde S; Aguila J; Zhong W; Solarz A; Mei I; Prud'homme J; Palkovits M; Turecki G; Mulder J; Uhlén M; Nagy C; Mechawar N; Hedlund E; Hökfelt T
Journal article: NEURON. 2023;111(23):3739-3741

Challenges of profiling motor neuron transcriptomes from human spinal cord.

Gautier O; Blum JA; Maksymetz J; Chen D; Schweingruber C; Mei I; Hermann A; Hackos DH; Hedlund E; Ravits J; Gitler AD
Article: METHODS IN MOLECULAR BIOLOGY. 2023;2594:143-164

Single-Cell mRNA-Seq of In Vitro-Derived Human Neurons Using Smart-Seq2.

Schweingruber C; Nijssen J; Benitez JA; Hedlund E
Article: SKELETAL MUSCLE. 2022;12(1):22

Mouse models of SMA show divergent patterns of neuronal vulnerability and resilience.

Woschitz V; Mei I; Hedlund E; Murray LM
Article: CELL METABOLISM. 2021;33(11):2215-2230.e8

Muscle-secreted neurturin couples myofiber oxidative metabolism and slow motor neuron identity.

Correia JC; Kelahmetoglu Y; Jannig PR; Schweingruber C; Shvaikovskaya D; Zhengye L; Cervenka I; Khan N; Stec M; Oliveira M; Nijssen J; Martínez-Redondo V; Ducommun S; Azzolini M; Lanner JT; Kleiner S; Hedlund E; Ruas JL
Article: NATURE MEDICINE. 2021;27(4):640-646

Altered perivascular fibroblast activity precedes ALS disease onset.

Månberg A; Skene N; Sanders F; Trusohamn M; Remnestål J; Szczepińska A; Aksoylu IS; Lönnerberg P; Ebarasi L; Wouters S; Lehmann M; Olofsson J; von Gohren Antequera I; Domaniku A; De Schaepdryver M; De Vocht J; Poesen K; Uhlén M; Anink J; Mijnsbergen C; Vergunst-Bosch H; Hübers A; Kläppe U; Rodriguez-Vieitez E; Gilthorpe JD; Hedlund E; Harris RA; Aronica E; Van Damme P; Ludolph A; Veldink J; Ingre C; Nilsson P; Lewandowski SA
Article: FRONTIERS IN MOLECULAR NEUROSCIENCE. 2021;14:699562

Spatial RNA Sequencing Identifies Robust Markers of Vulnerable and Resistant Human Midbrain Dopamine Neurons and Their Expression in Parkinson's Disease.

Aguila J; Cheng S; Kee N; Cao M; Wang M; Deng Q; Hedlund E
Article: GLIA. 2021;69(1):124-136

Disrupted function of lactate transporter MCT1, but not MCT4, in Schwann cells affects the maintenance of motor end-plate innervation.

Bouçanova F; Pollmeier G; Sandor K; Morado Urbina C; Nijssen J; Médard J-J; Bartesaghi L; Pellerin L; Svensson CI; Hedlund E; Chrast R
Article: NATURE COMMUNICATIONS. 2020;11(1):6341

Aberrant interaction of FUS with the U1 snRNA provides a molecular mechanism of FUS induced amyotrophic lateral sclerosis.

Jutzi D; Campagne S; Schmidt R; Reber S; Mechtersheimer J; Gypas F; Schweingruber C; Colombo M; von Schroetter C; Loughlin FE; Devoy A; Hedlund E; Zavolan M; Allain FH-T; Ruepp M-D
Article: GENOME RESEARCH. 2020;30(8):1083-1096

LCM-seq reveals unique transcriptional adaptation mechanisms of resistant neurons and identifies protective pathways in spinal muscular atrophy.

Nichterwitz S; Nijssen J; Storvall H; Schweingruber C; Comley LH; Allodi I; Lee MVD; Deng Q; Sandberg R; Hedlund E
Article: CELL REPORTS. 2020;31(9):107699

Radiation Triggers a Dynamic Sequence of Transient Microglial Alterations in Juvenile Brain.

Osman AM; Sun Y; Burns TC; He L; Kee N; Oliva-Vilarnau N; Alevyzaki A; Zhou K; Louhivuori L; Uhlén P; Hedlund E; Betsholtz C; Lauschke VM; Kele J; Blomgren K
Article: ACTA NEUROPATHOLOGICA. 2020;139(5):837-853

Synaptotagmin 13 is neuroprotective across motor neuron diseases.

Nizzardo M; Taiana M; Rizzo F; Aguila Benitez J; Nijssen J; Allodi I; Melzi V; Bresolin N; Comi GP; Hedlund E; Corti S
Article: BIO-PROTOCOL. 2019;9(14):e3312

Axon-seq for in Depth Analysis of the RNA Content of Neuronal Processes.

Nijssen J; Aguila J; Hedlund E
Article: STEM CELL REPORTS. 2019;12(6):1329-1341

Modeling Motor Neuron Resilience in ALS Using Stem Cells.

Allodi I; Nijssen J; Benitez JA; Schweingruber C; Fuchs A; Bonvicini G; Cao M; Kiehn O; Hedlund E
Article: JOURNAL OF PHYSIOLOGY-LONDON. 2019;597(12):3133-3146

Intact single muscle fibres from SOD1G93A amyotrophic lateral sclerosis mice display preserved specific force, fatigue resistance and training-like adaptations.

Cheng AJ; Allodi I; Chaillou T; Schlittler M; Ivarsson N; Lanner JT; Thams S; Hedlund E; Andersson DC
Article: HUMAN MOLECULAR GENETICS. 2019;28(10):1629-1644

Characterization of molecular mechanisms underlying the axonal Charcot-Marie-Tooth neuropathy caused by MORC2 mutations.

Sancho P; Bartesaghi L; Miossec O; García-García F; Ramírez-Jiménez L; Siddell A; Åkesson E; Hedlund E; Laššuthová P; Pascual-Pascual SI; Sevilla T; Kennerson M; Lupo V; Chrast R; Espinós C
Article: NATURE. 2019;567(7747):234-238

A radical switch in clonality reveals a stem cell niche in the epiphyseal growth plate.

Newton PT; Li L; Zhou B; Schweingruber C; Hovorakova M; Xie M; Sun X; Sandhow L; Artemov AV; Ivashkin E; Suter S; Dyachuk V; El Shahawy M; Gritli-Linde A; Bouderlique T; Petersen J; Mollbrink A; Lundeberg J; Enikolopov G; Qian H; Fried K; Kasper M; Hedlund E; Adameyko I; Sävendahl L; Chagin AS
Article: STEM CELL REPORTS. 2018;11(6):1565-1578

Axon-Seq Decodes the Motor Axon Transcriptome and Its Modulation in Response to ALS.

Nijssen J; Aguila J; Hoogstraaten R; Kee N; Hedlund E
Article: NATURE IMMUNOLOGY. 2018;19(5):435-7

Fatal demyelinating disease is induced by monocyte-derived macrophages in the absence of TGF-β signaling.

Lund H; Pieber M; Parsa R; Grommisch D; Ewing E; Kular L; Han J; Zhu K; Nijssen J; Hedlund E; Needhamsen M; Ruhrmann S; Guerreiro-Cacais AO; Berglund R; Forteza MJ; Ketelhuth DFJ; Butovsky O; Jagodic M; Zhang X-M; Harris RA
Article: MOLECULAR BIOLOGY OF THE CELL. 2018;29(2):75-83

CRISPR-Trap: a clean approach for the generation of gene knockouts and gene replacements in human cells.

Reber S; Mechtersheimer J; Nasif S; Benitez JA; Colombo M; Domanski M; Jutzi D; Hedlund E; Ruepp M-D
Article: METHODS IN MOLECULAR BIOLOGY. 2018;1649:95-110

LCM-Seq: A Method for Spatial Transcriptomic Profiling Using Laser Capture Microdissection Coupled with PolyA-Based RNA Sequencing.

Nichterwitz S; Benitez JA; Hoogstraaten R; Deng Q; Hedlund E
Article: MOLECULAR METABOLISM. 2018;7:12-22

Neurturin is a PGC-1α1-controlled myokine that promotes motor neuron recruitment and neuromuscular junction formation.

Mills R; Taylor-Weiner H; Correia JC; Agudelo LZ; Allodi I; Kolonelou C; Martinez-Redondo V; Ferreira DMS; Nichterwitz S; Comley LH; Lundin V; Hedlund E; Ruas JL; Teixeira AI
Article: STEM CELL REPORTS. 2017;9(3):742-751

Direct Reprogramming of Resident NG2 Glia into Neurons with Properties of Fast-Spiking Parvalbumin-Containing Interneurons.

Pereira M; Birtele M; Shrigley S; Benitez JA; Hedlund E; Parmar M; Ottosson DR
Article: GENOME RESEARCH. 2016;26(10):1342-1354

Single-cell analyses of X Chromosome inactivation dynamics and pluripotency during differentiation.

Chen G; Schell JP; Benitez JA; Petropoulos S; Yilmaz M; Reinius B; Alekseenko Z; Shi L; Hedlund E; Lanner F; Sandberg R; Deng Q
Article: NATURE COMMUNICATIONS. 2016;7:12139

Laser capture microscopy coupled with Smart-seq2 for precise spatial transcriptomic profiling.

Nichterwitz S; Chen G; Aguila Benitez J; Yilmaz M; Storvall H; Cao M; Sandberg R; Deng Q; Hedlund E
Article: SCIENTIFIC REPORTS. 2016;6:26448

Dopamine Receptor Antagonists Enhance Proliferation and Neurogenesis of Midbrain Lmx1a-expressing Progenitors.

Hedlund E; Belnoue L; Theofilopoulos S; Salto C; Bye C; Parish C; Deng Q; Kadkhodaei B; Ericson J; Arenas E; Perlmann T; Simon A
Article: SCIENTIFIC REPORTS. 2016;6:25960

Differential neuronal vulnerability identifies IGF-2 as a protective factor in ALS.

Allodi I; Comley L; Nichterwitz S; Nizzardo M; Simone C; Benitez JA; Cao M; Corti S; Hedlund E
Article: JOURNAL OF COMPARATIVE NEUROLOGY. 2016;524(7):1424-1442

Cross-disease comparison of amyotrophic lateral sclerosis and spinal muscular atrophy reveals conservation of selective vulnerability but differential neuromuscular junction pathology.

Comley LH; Nijssen J; Frost-Nylen J; Hedlund E
Article: ACTA NEUROPATHOLOGICA. 2016;131(3):453-464

Presymptomatic activation of the PDGF-CC pathway accelerates onset of ALS neurodegeneration.

Lewandowski SA; Nilsson I; Fredriksson L; Lönnerberg P; Muhl L; Zeitelhofer M; Adzemovic MZ; Nichterwitz S; Lawrence DA; Hedlund E; Eriksson U
Article: NEUROSCIENCE. 2015;291:216-229

Motor neurons with differential vulnerability to degeneration show distinct protein signatures in health and ALS.

Comley L; Allodi I; Nichterwitz S; Nizzardo M; Simone C; Corti S; Hedlund E
Article: STEM CELLS TRANSLATIONAL MEDICINE. 2014;3(9):1032-1042

Selection Based on FOXA2 Expression Is Not Sufficient to Enrich for Dopamine Neurons From Human Pluripotent Stem Cells.

Aguila JC; Blak A; van Arensbergen J; Sousa A; Vázquez N; Aduriz A; Gayosso M; Lopez Mato MP; Lopez de Maturana R; Hedlund E; Sonntag K-C; Sanchez-Pernaute R
Article: DEVELOPMENT. 2011;138(16):3399-3408

Specific and integrated roles of Lmx1a, Lmx1b and Phox2a in ventral midbrain development.

Deng Q; Andersson E; Hedlund E; Alekseenko Z; Coppola E; Panman L; Millonig JH; Brunet J-F; Ericson J; Perlmann T
Article: CELL STEM CELL. 2011;8(6):663-675

Transcription factor-induced lineage selection of stem-cell-derived neural progenitor cells.

Panman L; Andersson E; Alekseenko Z; Hedlund E; Kee N; Mong J; Uhde CW; Deng Q; Sandberg R; Stanton LW; Ericson J; Perlmann T
Article: BRAIN. 2010;133(Pt 8):2313-2330

Global gene expression profiling of somatic motor neuron populations with different vulnerability identify molecules and pathways of degeneration and protection.

Hedlund E; Karlsson M; Osborn T; Ludwig W; Isacson O
Article: STEM CELLS. 2008;26(6):1526-1536

Embryonic stem cell-derived Pitx3-enhanced green fluorescent protein midbrain dopamine neurons survive enrichment by fluorescence-activated cell sorting and function in an animal model of Parkinson's disease.

Hedlund E; Pruszak J; Lardaro T; Ludwig W; Viñuela A; Kim K-S; Isacson O
Article: PROCEEDINGS OF THE NATIONAL ACADEMY OF SCIENCES OF THE UNITED STATES OF AMERICA. 2008;105(15):5856-5861

Neurons derived from reprogrammed fibroblasts functionally integrate into the fetal brain and improve symptoms of rats with Parkinson's disease.

Wernig M; Zhao J-P; Pruszak J; Hedlund E; Fu D; Soldner F; Broccoli V; Constantine-Paton M; Isacson O; Jaenisch R
Article: STEM CELLS. 2007;25(5):1126-1135

Selection of embryonic stem cell-derived enhanced green fluorescent protein-positive dopamine neurons using the tyrosine hydroxylase promoter is confounded by reporter gene expression in immature cell populations.

Hedlund E; Pruszak J; Ferree A; Viñuela A; Hong S; Isacson O; Kim K-S
Article: NEUROSCIENCE. 2006;142(2):343-354

A tyrosine hydroxylase-yellow fluorescent protein knock-in reporter system labeling dopaminergic neurons reveals potential regulatory role for the first intron of the rodent tyrosine hydroxylase gene.

Kelly BB; Hedlund E; Kim C; Ishiguro H; Isacson O; Chikaraishi DM; Kim K-S; Feng G
Article: JOURNAL OF NEUROCHEMISTRY. 2006;97(5):1467-1480

Genetic selection of sox1GFP-expressing neural precursors removes residual tumorigenic pluripotent stem cells and attenuates tumor formation after transplantation.

Chung S; Shin B-S; Hedlund E; Pruszak J; Ferree A; Kang UJ; Isacson O; Kim K-S
Article: EXPERIMENTAL NEUROLOGY. 2005;194(2):363-375

L1 CAM expression is increased surrounding the lesion site in rats with complete spinal cord transection as neonates.

Kubasak MD; Hedlund E; Roy RR; Carpenter EM; Edgerton VR; Phelps PE
Article: MOLECULAR AND CELLULAR NEUROSCIENCE. 2005;28(2):241-252

The homeodomain transcription factor Pitx3 facilitates differentiation of mouse embryonic stem cells into AHD2-expressing dopaminergic neurons.

Chung S; Hedlund E; Hwang M; Kim DW; Shin B-S; Hwang D-Y; Kang UJ; Isacson O; Kim K-S
Article: EXPERIMENTAL NEUROLOGY. 2004;190(1):122-132

Region-specific cell grafting into cervical and lumbar spinal cord in rat: a qualitative and quantitative stereological study.

Kakinohana O; Cizkova D; Tomori Z; Hedlund E; Marsala S; Isacson O; Marsala M
Article: JOURNAL OF NEUROSCIENCE RESEARCH. 2004;75(3):307-319

Identification of a Hoxd10-regulated transcriptional network and combinatorial interactions with Hoxa10 during spinal cord development.

Hedlund E; Karsten SL; Kudo L; Geschwind DH; Carpenter EM
Article: MECHANISMS OF DEVELOPMENT. 2002;114(1-2):171-175

Differential Pax6 promoter activity and transcript expression during forebrain development.

Anderson TR; Hedlund E; Carpenter EM
Article: JOURNAL OF BIOLOGICAL CHEMISTRY. 2001;276(26):23937-23944

Neurosteroid hydroxylase CYP7B: vivid reporter activity in dentate gyrus of gene-targeted mice and abolition of a widespread pathway of steroid and oxysterol hydroxylation.

Rose K; Allan A; Gauldie S; Stapleton G; Dobbie L; Dott K; Martin C; Wang L; Hedlund E; Seckl JR; Gustafsson JA; Lathe R
Article: TRENDS IN PHARMACOLOGICAL SCIENCES. 1998;19(3):82-85

Cytochrome P450 in the brain: 2B or not 2B.

Hedlund E; Gustafsson JA; Warner M
Article: ARCHIVES OF TOXICOLOGY. SUPPLEMENT. ARCHIV F�R TOXIKOLOGIE. SUPPLEMENT. 1998;20:455-463

Extrahepatic cytochrome P450: role in in situ toxicity and cell-specific hormone sensitivity.

Warner M; Hellmold H; Magnusson M; Rylander T; Hedlund E; Gustafsson JA
Article: MOLECULAR PHARMACOLOGY. 1996;50(2):342-350

Cytochrome P4502D4 in the brain: specific neuronal regulation by clozapine and toluene.

Hedlund E; Wyss A; Kainu T; Backlund M; Köhler C; Pelto-Huikko M; Gustafsson JA; Warner M
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Preprint: BIORXIV. 2023

Human motor neurons are rare and can be transcriptomically divided into known subtypes

Gautier O; Blum J; Maksymetz J; Chen D; Schweingruber C; Mei I; Hermann A; Hackos D; Hedlund E; Ravits J; Gitler A
Preprint: BIORXIV. 2023

Single cell RNA sequencing in isogenicFUSandTARDBPmutant ALS lines reveals early mitochondrial dysfunction as a common pathway in motor neurons

Schweingruber C; Nijssen J; Mechtersheimer J; Reber S; Keuper M; Mei I; Benitez JA; O’Brien N; Jastroch M; Ruepp M-D; Hedlund E
Preprint: BIORXIV. 2022

Expression of substance P, NPY and their Receptors Is Altered in Major Depression

Barde S; Aguila J; Zhong W; Solarz A; Mei I; Prud’homme J; Palkovits M; Turecki G; Mulder J; Uhlén M; Nagy C; Mechawar N; Hedlund E; Hökfelt T
Review: BIOLOGY-BASEL. 2022;11(8):1191

The Cell Autonomous and Non-Cell Autonomous Aspects of Neuronal Vulnerability and Resilience in Amyotrophic Lateral Sclerosis.

Schweingruber C; Hedlund E
Corrigendum: NATURE MEDICINE. 2021;27(7):1308

Publisher Correction: Altered perivascular fibroblast activity precedes ALS disease onset.

Månberg A; Skene N; Sanders F; Trusohamn M; Remnestål J; Szczepińska A; Aksoylu IS; Lönnerberg P; Ebarasi L; Wouters S; Lehmann M; Olofsson J; von Gohren Antequera I; Domaniku A; De Schaepdryver M; De Vocht J; Poesen K; Uhlén M; Anink J; Mijnsbergen C; Vergunst-Bosch H; Hübers A; Kläppe U; Rodriguez-Vieitez E; Gilthorpe JD; Hedlund E; Harris RA; Aronica E; Van Damme P; Ludolph A; Veldink J; Ingre C; Nilsson P; Lewandowski SA
Preprint: BIORXIV. 2021

Muscle-secreted neurturin couples myofiber oxidative metabolism and slow motor neuron identity

Correia JC; Kelahmetoglu Y; Jannig PR; Schweingruber C; Svaikovskaya D; Zhengye L; Cervenka I; Oliveira M; Nijssen J; Martínez-Redondo V; Stec M; Khan N; Lanner JT; Kleiner S; Hedlund E; Ruas JL
Preprint: BIORXIV. 2020

LCM-seq identifies robust markers of vulnerable and resistant human midbrain dopamine neurons

Aguila J; Cheng S; Kee N; Cao M; Deng Q; Hedlund E
Conference publication: EUROPEAN JOURNAL OF HUMAN GENETICS. 2019;27:1479-1480

Expanding the molecular characterization of the CMT2Z disease associated gene MORC2

Sancho P; Bartesaghi L; Moissec O; Garcia-Garcia F; Ramirez-Jimenez L; Siddell A; Akesson E; Hedlund E; Lassuthova P; Pascual-Pascual S; Sevilla T; Kennerson M; Lupo V; Chrast R; Espinos C
Preprint: BIORXIV. 2018

Modeling motor neuron resilience in ALS using stem cells

Allodi I; Nijssen J; Benitez JA; Schweingruber C; Fuchs A; Bonvicini G; Cao M; Kiehn O; Hedlund E
Preprint: BIORXIV. 2018

LCM-seq reveals unique transcriptional adaption mechanisms of resistant neurons in spinal muscular atrophy

Nichterwitz S; Storvall H; Nijssen J; Comley LH; Allodi I; van der Lee M; Schweingruber C; Deng Q; Sandberg R; Hedlund E
Review: MOLECULAR ASPECTS OF MEDICINE. 2018;59:36-46

Single-cell RNA sequencing: Technical advancements and biological applications.

Hedlund E; Deng Q
Review: ACTA NEUROPATHOLOGICA. 2017;133(6):863-885

Motor neuron vulnerability and resistance in amyotrophic lateral sclerosis.

Nijssen J; Comley LH; Hedlund E
Conference publication: NEUROLOGY. 2017;88

The oculomotor restricted protein synaptotagmin 13 protects motor neurons from degeneration in ALS and SMA

Nizzardo M; Rizzo F; Taiana M; Allodi I; Benitez JA; Nijssen J; Ulzi G; Melzi V; Del Bo R; Bresolin N; Comi G; Hedlund E; Corti S
Conference publication: ACTA PHYSIOLOGICA. 2017;219:17

Increased fatigue resistance and preserved specific force in intact single muscle fibres from the SOD1G93A mouse model of ALS

Cheng A; Allodi I; Chaillou T; Thams S; Ivarsson N; Schlittler M; Lanner J; Hedlund E; Andersson D
Review: CELLULAR AND MOLECULAR LIFE SCIENCES. 2014;71(6):999-1015

Cellular therapy to target neuroinflammation in amyotrophic lateral sclerosis.

Rizzo F; Riboldi G; Salani S; Nizzardo M; Simone C; Corti S; Hedlund E
Review: FRONTIERS IN NEUROSCIENCE. 2014;8:109

Directed midbrain and spinal cord neurogenesis from pluripotent stem cells to model development and disease in a dish.

Allodi I; Hedlund E
Review: STEM CELLS INTERNATIONAL. 2012;2012:412040-17

Cellular programming and reprogramming: sculpting cell fate for the production of dopamine neurons for cell therapy.

Aguila JC; Hedlund E; Sanchez-Pernaute R
Editorial comment: EXPERIMENTAL NEUROLOGY. 2011;231(1):14-18

The protective effects of β-lactam antibiotics in motor neuron disorders.

Hedlund E
Review: JOURNAL OF INTERNAL MEDICINE. 2009;266(4):358-371

Neuronal cell replacement in Parkinson's disease.

Hedlund E; Perlmann T
Meeting abstract: MECHANISMS OF DEVELOPMENT. 2009;126:S245

Intrinsic transcriptional determinants promote efficient generation of neuronal subtypes from ES cells

Panman L; Andersson E; Hedlund E; Udhe C; Mong J; Alexsenko Z; Sandberg R; Ericson J; Perlmann T
Editorial comment: CELL STEM CELL. 2008;3(6):575-576

ALS model glia can mediate toxicity to motor neurons derived from human embryonic stem cells.

Hedlund E; Isacson O
Review: EUROPEAN JOURNAL OF NEUROSCIENCE. 2007;26(7):1721-1737

Cell therapy and stem cells in animal models of motor neuron disorders.

Hedlund E; Hefferan MP; Marsala M; Isacson O
Review: CURRENT DRUG METABOLISM. 2001;2(3):245-263

Cytochrome P450 in the brain; a review.

Hedlund E; Gustafsson JA; Warner M
Published conference paper: MUTATION RESEARCH-FUNDAMENTAL AND MOLECULAR MECHANISMS OF MUTAGENESIS. 1997;376(1-2):79-85

Cytochrome P450 in the breast and brain: role in tissue-specific activation of xenobiotics.

Warner M; Hellmold H; Yoshida S; Liao D; Hedlund E; Gustafsson JA
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Anställningar

Senior Forskare, Cell- och molekylärbiologi, Karolinska Institutet, 2023-

Examina och utbildning

Docent, neurobiologi, karolinska institutet, 2011
MEDICINE DOKTORSEXAMEN, Inst medicinsk näringslära - nedl 060101, Karolinska Institutet, 2000

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